1912] EAST—HYBRIDS OF NICOTIANA 247 
rich ground the following year.4 Each plant among the progeny 
had numerous three-celled capsules, together with occasional two- 
celled and four-celled capsules. For two generations the strain has 
bred true to this condition. The only other abnormality observed 
is the occurrence of a greater number of flowers with six sepals and 
six petals than is common in the normal NV. Bigelovii or the NV. 
quadrivalvis. About 1 per cent of the flowers from the normal 
species have the extra petal and sepal, but on individual plants in 
Fic. 4.—At left, Nicotiana Bigelovii var. quadrivalvis; at right, N. quadrivalvis 
Pursh; in center, F, generation of cross. 
the aberrant strain of N. Bigelovii from 2 to 5 per cent of the 
flowers vary in this manner. 
The aberrant N. Bigelovii was crossed reciprocally with N. 
quadrivalvis. The F, plants were: alike in each case. They were 
intermediate in stature and in size of flower. The earlier capsules 
were four-celled; later in the season both three cells and two cells 
were produced. The F, plants were fully as fertile as the parent 
species. Each plant produced hundreds of well filled capsules. 
4 Seeds are always started in sterilized soil and seedlings set in the open. 
