248 BOTANICAL GAZETTE [MARCH 
One plant each from the cross and its reciprocal were selfed. About 
zo plants were grown from each mother plant. No difference was 
noticed in the two F, generations. The entire lot can therefore be 
considered together. The plants varied in height from one foot to 
two feet. The flowers were in general intermediate in size, but 
varied to the extremes that characterize each parent. No plants 
having only two-celled capsules were found; 71 had a large number 
of four-celled capsules; 40 had only a few four-celled capsules; 
while 19 had no four-celled capsules. Every plant had large 
numbers of three-celled and two-celled capsules, whether or not 
four-celled capsules were present. 
Summary 
1. Two elementary species of N. Bigelovii Pursh have been 
found. In one the capsules are two-celled and selection of indi- 
viduals having an occasional three-celled capsule does not increase 
the tendency; in the other the tendency to have a greater number 
of cells than two in the capsule is always transmitted. 
2. N. quadrivalvis Pursh and normal NV. Bigelovii Watson are 
alike in all specific characters except the number of cells in the 
capsule, and since they give fertile hybrids when crossed it is thought 
that NV. Bigelovii gave rise to N. quadrivalvis. 
3. It is proposed that the section Potrrcita in the genus Nico- 
tiana be dropped, and N. quadrivalvis Pursh be called N. Bigelovit 
var. guadrivalvis. 
Harvarp UNIVERSITY. 
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