204 BOTANICAL GAZETTE [APRIL 
It was also noted above that no split in the chromosomes is 
evident in the anaphase of the first division. But though such a 
split is stated by GriGorRE to be a part of his ‘‘scheme,” its absence 
cannot be regarded as a very important deviation. 
THE CHROMOSOMES IN THE REDUCTION DIVISION 
As the chromosomes become vacuolate during interkinesis, the 
stage most favorable for an examination and comparison of the 
chromosomes is the anaphase of the reduction division (fig. 13). 
The most striking thing about the chromosomes at this stage is 
their different size in the two forms. The chromosomes of the 
short-styled form have a diameter nearly twice as great as do those 
of the long-styled form (compare figs. 14 and 15 with figs. 16 and 
17). As it is entirely improbable that this difference in size can 
be due to any difference in the hereditary qualities borne by the 
two sets of chromosomes, it appears to be related with the corre- 
sponding, though smaller, difference in the size of the microspore 
mother cells at this stage. 
Just what relation exists between the size of the cell, the size of 
the nucleus, and the size and mass of the chromosomes is not well 
understood. There seems, at any rate, to be no definite relation 
between the number of chromosomes and the size of the nucleus. 
STRASBURGER (36, p. 51) cites several cases in which the nuclei 
with the diploid number of chromosomes are distinctly larger than 
those with the haploid number in the same species. In the seed 
rudiments of Taxus baccata, for example, the nuclei of the pro- 
thallium are much smaller than those of the nucellus. That the 
diploid number of chromosomes is not always associated with a 
larger nucleus than that which contains the haploid number, how- 
ever, he proves by the case of Dictyota dichotoma, in which species the 
nuclei of the plants which produce tetraspores are no larger than 
those of the plants which bear eggs or sperms, though they have, 
of course, twice as many chromosomes. 
Another difference between the two forms, which is apparently 
constant, is the arrangement of the chromosomes in the anaphase 
of the heterotypic division. In the short-styled form the eight 
chromosomes tend to be arranged with six in the peripheral ring 
