498 BOTANICAL GAZETTE [JUNE 
studied here, nor do either CAMPBELL (2) or LeITGEB (1) report such 
a condition. 
True elaters are present, long, slender, fusiform, and thickened 
by usually two spiral bands. 
The spores themselves have fairly thick walls, highly sculptured, 
and are produced in great numbers (fig. 11). CAVERS (3) has 
described the spore mother cells as lobing 
deeply before division, as happens in many 
of the Jungermanniales. All of the prepara- 
tions studied here, however, show the ordi- 
nary tetrad formed from a spherical spore 
mother cell. 
Seta and foot.—Both seta and foot are 
rather well developed. While small as com- 
pared with the size of the capsule, they are 
nevertheless rather massive, and well differentiated from each 
other. The foot is club- 
shaped, and bluntly OS tiie ea 
pointed at the lower end 
(fig. 12). Thesurface cells 
project as short papillae, 
Fic. 11.—Tuberculate 
spore. 
Seek : — 
and are haustorial in func = a5ea\ 
tion. These cells, as well Se came BS ae 
See aT 
as those of the calyptra KE BR) 
° i . ehA- Oe! 
surrounding them, show in A ip SY 
section a very much darker 
stain than do the cells of 
the surrounding tissues. 
Between the calyptra and 
foot is interposed a fairly Bd 
thick layer of mucilaginous Fic. 12.—Foot and seta 
material. 
Although 6-10 archegonia are produced, only one egg is fertil- 
ized. The remaining archegonia, however, persist, and are still 
visible after the sporophyte has matured; soon after the sporophyte 
has begun its development, however, their contents break down 
into a darkly staining mucilaginous mass. 
