222 BOTANICAL GAZETTE [SEPTEMBER - 
Hutton (15) to Thuyitis because of the verticillate arrangement of 
the leaves. SCHIMPER (22) then transferred them to STERNBERG’S 
taxodineous genus Echinostrobus, where the leaves are sometimes 
spiral and sometimes whorled. Saporta (20), however, pointed 
out the inadvisability of this step and put them back into the 
Cupressineae, this time as Palaeocyparis expansa. SEWARD (24) 
refers to the difficulty of distinguishing between the whorled leaves 
of Thuyitis and the spiral ones of Brachyphyllum, and suggests (23) 
that, at least so far as the specimens of LINDLEY and HUTTON are 
concerned, the two genera are identical. The structure of the 
cuticle is not without bearing on this question, for in the species 
of the latter which have been examined—B. macrocarpum NEw- 
BURY (14), B. Miinsteri, and B. affine (21)—the stomata are in 
long rows alternating with strands of sclerenchyma. ‘This condi- 
tion, of course, is entirely different from that of Thuyitis expansus, 
though it is singularly like 7. Schlonbachi ScHENK (21) and the 
living podocarpineous genus Microcachrys. For a parallel, we are 
driven to the Taxodineae (Arthrotaxis) or the Cupressineae (Thuja, 
etc.), and though, as suggested above in the case of Palissya, 
phyllotaxy is not an invariable test for affinities, still in view of 
the fact that, as a whole, the leaves of the Taxodineae are in spirals 
and those of the Cupressineae in whorls, it would seem to be 
advisable to retain the original name, and, at least until an examina- 
tion of their internal structure settles the question of affinities, to 
continue to call shoots of this type Thuyitis expansus. 
i 
Taxitis tenerrimus 
The next specimen to be described has been referred by FEIsT- 
MANTEL (10) to Tavitis tenerrimus, and the spiral arrangement. of 
the linear, uninerved, and decurrent leaves, shown in fig. 3, indi- 
cates the correctness of this identification. The cuticle of the 
upper surface is entirely devoid of stomata; that of the lower is 
represented in fig. 8. The epidermal cells are irregular in shape, 
with a slight tendency to become elongated below the midrib. 
The stomata are scattered without definite arrangement, but the 
indifferent state of preservation prevents any detailed description. 
In general, however, there are 4—6 accessory cells, and the opening 
