Igt5] AASE—M EGASPOROPHYLLS OF CONIFERS 309 
In the second group based on vascular anatomy the bract and 
scale vascular supplies are more or less intimately united into one 
bundle which springs from the base of the cylinder gap. 
Of sporophylls with parts separate there belong here the lower 
sporophylls of Pinus Banksiana, P. maritima, and Keteleeria 
Fortunei (the ovules in these sporophylls are to greater or less 
extent abortive), and Cedrus Libani (2). Of the sporophylls with 
parts considerably united there fall into this group those of Cun- 
ninghamia, Arthrotaxis laxifolia (1), most species of Arauéaria 
(1, 3, 6, '7, 9, 10), and the lower sporophylls of Cryptomeria japonica, 
Cupressus Benthamii, and Thuja occidentalis. Most of the appar- 
ently simple sporophylls are included in this group, as those of 
Agathis, Saxegothaea (5, 8), and Arthrotaxis selaginoides (1). 
The degree of welding of the bract and scale vascular supplies 
varies considerably. In Arthrotaxis selaginoides, Agathis, Arau- 
caria, and Saxegothaea the two remain united into one bundle for 
greater or less distance in the cortex; in most of the others the single 
bundle divides early; but in many cases, where the two sporophy]l 
parts have fused extensively, branches of the scale supply swing 
about to lie on the ventral side of the appendage at each side of the 
bract bundle. This fact is well illustrated in Thuja occidentalis, 
Juniperus communis, Cupressus Benthamii, and Cryptomeria 
japonica. In Cupressus Benthamii and Cryptomeria japonica the 
scale bundles at either side of the bract bundle even accompany 
the bract bundle into the free portion of the bract. 
Bundle distribution is generally directly related to the size of the 
organ supplied, hence the bundles extend into the most expanded 
region of the-sporophyll, whether that particular region represents 
bract or scale. 
In Cunninghamia, Araucaria, and Agathis, in which absence 
of sporophylls with separate bract and scale supplies makes com- 
parison impossible, it is difficult to determine with certainty what 
is bract and what is scale supply. The matter is further compli- 
cated by the presence in the last two genera of a branching bundle 
in the vegetative leaf, a condition which probably implies a branch- 
ing bract bundle in the bract of the sporophyll as well. And, 
further, Cupressus Benthamii and Cryptomeria have clearly shown 
