312 BOTANICAL GAZETTE [OCTOBER 
3. Simplification of a compound sporophyll has been attained 
to fullest extent in Arthrotaxis selaginoides, Agathis, and Saxego- 
thaea, and possibly others; an extensive reduction of bract occurs 
in Cedrus Libani and the lower sporophylls of Pinus maritima; the 
scale in Phyllocladus is probably reduced so as to be represented 
only by a distinct ovular supply; the welding of the two organs is 
complete in Juniperus communis and Chamaecyparis Lawsoniana. 
4. Fusion of bract and scale vascular supplies does not directly 
parallel fusion of bract and scale. 
5. Separate origin of bract and scale vascular supplies occurs 
most generally in the Podocarpineae and Abietineae; fusion of 
bract and scale supplies has reached its highest expression in the 
Araucarineae; both types of bundle origin are represented in the 
same strobilus in Cryptomeria japonica, Cupressus Benthamii, and 
the lower sporophylls of Pinus. 
6. The bract bundle in plants with uninerved vegetative leaves 
divides only slightly if at all; the extent of the scale bundle system 
is directly related to the size of the organ supplied. 
7. The scale bundles in the Abietineae and Chamaecyparis 
Lawsoniana form in the expanded portion of the organ a straight 
row or arc; in members of the Taxodineae and Cupressineae scale 
bundles swing around so as to lie at each side of the bract bundle. 
8. In Crypiomeria japonica and Cupressus Benthamii and per- 
haps Cunninghamia Davidiana scale bundles accompany the bract 
bundle into the free portion of the bract: 
g. A branching bundle in the vegetative leaf in Araucaria and 
Agathis probably implies a branching bundle in the bract of the 
sporophyll; the vascular system in the megasporophyll is probably 
a complex of bract and scale bundles. 
ro. In species of Podocarpus the scale bundles continue in the 
portion of the scale folded toward the dorsal side, forming the 
epimatium of the ovule. 
The author is indebted to Professor J. M. Courter and Dr. — 
C. J. CHAMBERLAIN for valuable suggestions, and to the latter also 
for excellent material. 
UNIVERSITY OF CHICAGO 
