370 BOTANICAL GAZETTE [NOVEMBER 
fall after blooming, but in some varieties they persist even after 
the ripening of the fruit. 
The staminate flower begins to develop as early as the pistillate, 
but the pistils develop more rapidly than the stamens, resulting 
in the sterilization of stamens and in some cases in their disap- 
pearance. 
The mature pistillate flower generally has two sets of stamens 
(nos. 1 and 2), but in some cases has more (nos. 1, 2, and some of 
4) or less (no. 1 and some of no. 2). 
I did not discover the following condition described by Miss 
Hacue: “In the pistillate flower it is a common occurrence to 
find the number increased by the branching of one or more of the 
stamens.” 
In the development of the pistil there appear first four large 
protrusions, and then four smaller ones between them (figs. 5 and 
6, a,b). The larger ones give rise to two small papillae, which are 
the initials of ovules (figs. 6, d, and 7). The papillae grow at first 
toward the wall of the ovary, then curve downward toward the 
center, and finally upward (figs. 8 and 9). Simultaneously the 
small protrusions develop toward the center of the ovary, resulting, 
along with the development of larger protrusions, in 8 loculi for the 
8 ovules. 
The ovule has two integuments (fig. 9), which extend beyond the 
nucellus like a beak at the time the embryo sac is fully developed 
(fig. 10). ; 
Megaspore formation 
In all the varieties examined the archesporial cell is solitary 
and hypodermal. There is no division resulting in a parietal cell, 
so that the archesporial cell is the megaspore mother cell. The 
heterotypic division of this nucleus occurs in the same way as in 
the case of the nucleus of the microspore mother cell. After 
nuclear division, the cytoplasm organizes into two parts, the inner 
one being the larger. Sometimes a wall between these two parts 
does not appear before another division of nuclei occurs. | 
Wall formation in connection with the megaspores is variable 
in direction. Sometimes the two walls are perpendicular to one 
