444 BOTANICAL GAZETTE [DECEMBER 
obtained from 15 capsules of the primary mutation, Lexington E-5- 
238. The progeny consisted of 25 plants, only 16 of which survived 
transplanting from the seed pan. None of the progeny resembled 
the parent. All were extreme dwarfs which resembled, but were 
not identical with, mut. revoluta and mut. setacea. They differed 
mainly in the thicker leaves, which in 4 plants were narrow but 
not markedly revolute. Although a very nondescript lot, differing 
much among themselves, 5 most resembled mut. revoluta, and 7 
mut. sefacea. The result of this culture might almost have been 
predicted. The mass mutability was inherited by mut. gigas from 
f. typica. In view of the dependence of the gigas characters upon 
TABLE V 
ANALYSIS OF AN F; CULTURE FROM SELF-POLLINATED MUT. Jatifolia, LEXINGTON E-36 
The parent plant belonged to the F, from Lexington E, f. typica; see table I, culture 2, 
for position in pedigree 
eae sokg ia | st 
Parent Seeds | Total eo 3 3 $ 3 3 3 $3 
planted} plants Z . = br be ne | ae ee 
5 3 3 3 3 | 26 | $2)| §8 
= = = = >on = a 
Lex. E-36, mut. 
latifolia. ....... aye" | 3827 65 | 4B) Ss 2 29 | 3t 87 | 47.8 
* Seeds from 7 capsules, containing respectively 54, 72, 49, 3°, 38, 79, am ‘inde 
t Nos. 32 and 43, a new mutation; no. 51 mut. gigas (?), morphologically identical with Lexington 
Esaas which had 28 chromosomes; see table II, culture 5 from Lexington E-5, for position of latter 
» 4 Eictading, of course, mut. /Jatifolia. 
the double complement of chromosomes, which would in general 
be handed on to any secondary mutations, it follows that the muta- 
tions occurring en masse would not be identical with those from 
f. typica. Furthermore, chance irregularities in chromosome 
distribution might increase the polymorphism of the progeny. 
In such a highly modified germ plasm irregularities would be 
expected. 
The cross mut. gigasXf. typica yielded 160 seeds in a single 
capsule, of which 10 germinated. The plants were all extreme 
dwarfs, of the most nondescript nature, hardly any two alike. All 
had thick leaves, some plane, others revolute. No mutation of 
