200 THE LORANTHACEAE OF AUSTRALIA, ii., 



which ai'e peripheral, and situated opposite the largest vascular bundles. The 

 peripheral canals pass into the midrib of the leaf with the vascular bundles, 

 retaining their position on the upper side of the latter; they do not appear to 

 undergo any further branching; but become considerably swollen at certain 

 points. The mucilage-canals of Nuytsia are provided with an epithelium of 

 several layers, but are nevertheless of lysigenous origin; in the leaf the middle 

 lamellae of the gelatinized cells, which fonn the canal, can still be clearly recog- 

 nised." t 



KEY TO THE SUBFAMILIES AND TRIBES. 



(A). Receptacle with "Calyculus," or a calyx-like outgrowth surrounding the 

 base of the perianth. Perianth double. Flowers bisexual. Filaments conspicu- 

 ous; anthers adnate or versatile, two-celled, the cells opening longitudinally. Fruit 



with a central layer of viscin surrounding the endosperm 



Subfam. I. LORANTHOIDEAE. 



Stems with mucilage canals in the pith, andj in the latter stages in the bast. 

 Calyculus rudimentary. Fruit dry, tripterous; endosperm soft, albuminous- Viscin 

 layer thin. Terrestrial root-parasitic shrubs or trees 1. NUYTSIEAE 



Stems without mucilage canals. Bast only outside of the wood. Calyculus 

 more or less developed. Fruit berry-like or like a stone-fruit. 2. LORANTHEAE. 



Ovary one or more celled (one-celled in all the Austrahan genera) ; embryo 

 sac only on fertilization pressing right up to the base of the style. Fruit drup- 

 aceous. Endosperm horny, deeply cleft. Viscin scanty. Terrestrial ? non-root 

 parasitic shrubs 2a. GAIADENDRINAE. 



Perfect ovary one-celled; embryo sac pressing up towards the base of the 

 style. Endosperm soft, albuminous, smooth, or with faint longitudinal depres- 

 sions. Viscin copious- Semi-parasitic shrubs 2b. LORANTHINAE. 



(B). Receptacle without "Calyculus," or a calyx-like outgrowth surrounding 

 the base of the perianth, or the perianth single. Flowers always unisexual. 

 Anthers sessile or nearly so, with three or numerous pollen-bearing chambers, 

 dehiscent by pores. Fruit with a sticky central layer. Seeds compressed in all 

 the Australian species Subfam. II- VISCOIDEAE. 



Placenta central. Anthers imperfectly 2-celled or dehiscent by a terminal 

 orifice. Perianth 3-merous 1. KORTHALSELLINEAE. 



Placenta basal. Anthers adnate to the petals, many chambered, dehiscent by 

 pores. Perianth 2— 4-merous ' 2a- VISCEAE. 



I. 1. LOEANTHOIDEAE-NUYTSIEAE. 



Engler in Engl, et Prantl, Pflanzenfam., iii., i., 1889, 177, et Nachtr., iii., 

 1897, 12i.—Euloranthsae Benth., in Bth. et Hook, f., Gen. PI., iii., 1880, 205. 



Calyculus rudimentary, shortly triquetrous. Fruit tripterous. Ovary 1- 

 celled. Cotyledons 2-4. Endocarp soft, not furrowed. Xylem with bands of 

 soft bast, traversed by concentric layers of thin-walled tissue, which consists 

 chiefly of parenchymatous cells and includes mucilage-canals, also small groups 

 of soft bast. 



1. Nuytsia R.Br. 



Journ. Geogr. Soc, i., 1831, 17; Benth., B.Fl., iii., 1866, 387. 



Flowers bisexual. Perianth double. Calyx shortly winged, the tube adnate 

 to the ovary; the limb short, irregularly 6-toothed. Petals 6, linear, free, erect 

 or spreading. Stamens as many as petals, inserted on the lower half of the 



t Solereder, Systematic Anatomy of the Dicotyledons, ii., 1908, p. 728. 



