﻿BOTANICAL GAZETTE 



the secondary one arises could be readily located. The landmark 

 selected is at the right in fig. 56, where there is a primary lamella 

 and at its left a secondary one quite well developed but close to it. 

 Following this in the successive figures up to fig. 61, it will be seen 

 that the two become closer and closer, until both have the same 

 trama at their junction with the pileus and appear like a double 

 lamella, since in this region the secondary lamella evidently sprouted 

 out of the base of the primary one. Now the area which we wish 

 to observe for the origin of a secondary lamella, as the primary 

 ones diverge more and more, is between the second and third 

 primary lamella to the left of this "double" lamella. This space 

 is between the two primary lamellae at the left of the figure. The 

 secondary lamella originates as a salient from the palisade layer of 

 the hymenophore between the two primary lamellae. In fig. 57 

 this space is broader. In fig. 58 there is seen a slight salient, some 

 of the marginal cells of which have swollen into cystidia. Then 

 in figs. 59, 60, and 61 it is more and more pronounced, showing 

 the same features as the primary lamellae, but is much narrower, 

 and because of the greater width of the primary lamellae its margin 

 is held away from the fundamental plectenchyma below, except in 

 figs. 60 and 61, where it is coming in contact with a few loose threads. 



The lack of fundamental plectenchyma in the trama of 

 the lamellae. — If the lamellae originated as described for C. mica- 

 ceus (Levine 22), by the splitting of primary structures which form 

 a series of radiating ridges isolated by fundamental plectenchyma or 

 elements, then, as the approximate halves of two adjacent ridges 

 turn toward each other to form a lamella, they would inclose some 

 of these fundamental elements between the pileus trama and the 

 trama of the lamella. There would be small islands of funda- 

 mental plectenchyma extending along the entire length in the base 

 of each lamella. This is not the case. On the other hand, all the 

 evidence goes to show that the trama is newly formed tissue and 

 grows downward from the trama of the pileus into the lamella, and 

 this is evident from the earliest origin of the first salients or ridges, 

 the trama lying in the ridges, not between them. For this reason 

 we may find, when we come to study carefully the origin of the 

 lamellae in Amanita and Amanitopsis, that the method of origin is not 



