﻿BOTANICAL GAZETTE 



described. A spreading or fan-tailing of the margins of the lamellae 

 immediately takes place, largely due to a swelling of the marginal 

 palisade into cystidia-like structures. It is this fan-tailing of the 

 gills, accompanied by the swelling of the marginal palisade cells 

 of the young salients into cystidia-like structures, which leads to 

 the situation observed by Levine (22, p. 351) in his fig. 8, of which 

 he says that the palisade cells do not inclose the edge of a gill, but 

 form an arched palisade layer in each "gill chamber" (22, p. 356). 

 Exactly such a situation is shown in figs. 56-63 of the present 

 article. The palisade layer no longer is present over the margins 

 of these gills because at an early stage of the gill origins the palisade 

 cells inclosing the edges of the salients become swollen into cystidia. 

 Figs. 8, 13, and 14 of Levine's paper represent quite old stages in 

 the development of the gills. 



Because of the weak annular gill cavity, the broadening of the 

 lamellae by growth soon brings them in closer contact with the stem, 

 or the rather thin layer of fundamental plectenchyma which clothes 

 the stem. Because of the loose and shredded character of this 

 fundamental plectenchyma, the loose hyphae readily interlock 

 with the swollen marginal cells of the lamellae, and with isolated 

 hyphae which grow down from the marginal trama of the lamellae. 

 The frazzled layer of fundamental plectenchyma on the surface of 

 the stem is much thinner in C. micaceus than in C. atramentarius . 

 For this reason the round or angular cells of the stem surface very 

 soon come in contact with the trama of the lamellae as the surface 

 cells are spread laterally by the increasing pressure of contact. 

 There is, therefore, a wedging together, to a greater or less extent, 

 of the trama and stem cells which often presents the appearance 

 of pseudoparenchyma. The different stages are represented in 

 figs. 55-63 and 65-67. 



The attachment of the lamellae to the stem, therefore, is 

 a gradual process, proceeding from the young to the older stages 

 of the lamellae and basidiocarp. It is interesting to note that 

 Levine's fig. 8 represents the weak annular gill cavity, below the 

 margins of the gills, which are loosely connected by isolated hyphae 

 or loose strands extending across the cavity to the stem, or the layer 

 of fundamental plectenchyma on its surface. It represents very 



