272 BOTANICAL GAZETTE [OCTOBER 
was counted in July, during the beginning of the flowering period, 
but about one-half of some of the cultures were kept in the boxes 
so as to reach only the stage of rosettes at that time. No second 
generations have been cultivated. I will first describe the results, 
and afterwards give the necessary details concerning the several 
cultures. 
The main purpose of these crosses was to decide the question 
whether the special characters of O. cana are handed down by the 
ovules only, as in O. scintillans and O. lata, or by the pollen also. 
Table IX gives the result of the crosses with wild species other 
than O. Lamarckiana. Here a splitting occurs into Jaefa and 
velutina or into densa and laxa, exactly analogous to that produced 
by O. Lamarckiana itself. The only exception is the pollen of 
O. biennis, which does not split; it gives with O. cana the same 
type as with the parent species. The result was very striking on 
the beds. No specimens of the cana type occurred in the cases 
where this mutant had been the father, whereas such individuals 
abounded in the results of the reciprocal crosses. 
TABLE IX 
CROSSES OF O. cana WITH OTHER SPECIES 
a ee | tee | ee | ee 
O. canaXO. beads Chicago 25 II 63 I 
O. canaXO. Cocke 17 17 63 3 
O. cana XO. Hookeri....... 28 14 66 oe lever vena 
O. Hookeri XO. cana,.:.. 2. ° 4 O0° 2 fs eee 
O. Cockerelli XO. cana... .. ° 54 MM hie 
O. syrticolaXO. cana...... ° 51 ee ey 
O. biennis XO. cana....... ° 58 ee 
he hag of — of 
O. biennis Chicago. cana ° 24 75 I 
O. atrovirens XO. c ° 36 ee 
Cc Z 
O. canaXO. biennis....... 49 Oe Oe hee EN On, 
In this table, O. syrticola Bartlett is the O. muricata L. of my 
Gruppenweise Artbildung, and O. atrovirens Bartlett has been 
