394 BOTANICAL GAZETTE [NOVEMBER 
of the blade. The marginal ones are larger. Inside this whorl of 
petals there are 5 stamens. These appear to be in every respect 
normal and compare favorably in size and shape with the stamens 
in normal flowers. Normal tetrad pollen grains have been found 
in their anthers (fig. 4). In the case (fig. 5) of one flower which is 
mentioned later, the filaments were broader and the anthers in some 
cases consisted of a single locule each, yet all intermediate stages 
between unilocular and bilocular anthers were present. In these 
anthers no pollen was found. Unlike PLANCcHON’s (16) chloranthic 
flowers of D. intermedia, these had no carpels or any indication of 
carpels (figs. 6,13). The center of the flower was occupied by 
a cluster of leaf rudiments in various stages of development, as 
described by FERNALD (3) and Leavitt (11) for D. rotundifolia 
leaf. The outer leaf rudiments are best developed, while the ones 
toward the center are progressively smaller (fig. 6). In older flowers 
these rudiments have become fully developed spatulate blades resem- 
bling those of the normal vegetative leaf. In a single capitate inflor- 
escence all stages in the development of the central leaf rudiments 
can be found. Fig. 7 represents a small chloranthic flower in such a 
head. The sepals are fully developed, while the petals, although 
they are fully expanded, have not all extended their tentacles. The 
unfolding leaflike petals show the characteristic incurved form 
common to the vegetative leaves. The stamens in this flower 
(fig. 7) are normal and are few in number. The carpels as such 
are entirely lacking, but are replaced by a small number of very 
minute leaf rudiments, of which one is considerably larger than the 
others. Figs. 8 and 9 represent larger chloranthic flowers. The 
sepals shown in fig. 8 are larger and much broader than the average 
I observed. In the center of the flower there is a large rosette of 
young leaves entirely replacing the carpels (fig. 8). The outer 
leaf rudiments are large and almost ready to unfold. Fig. 9 
represents a chloranthic flower in which the leaves are slightly 
farther advanced. Fig. ro represents a chloranthic flower in which 
the central leaves have attained the greatest development I found. 
The flower measured approximately 6 mm. in diameter. Only one 
petal and sepal are figured, but the usual number 5 was present 
here also. The outer whorl of the apical leaf group has fully 
