REPORT ON THE CIRRIPEDIA. If' 



chitinous outer wall is easily observed. The external surface of the body-wall is clothed 

 with microscopic spines, having a length of about 0'0235 mm., and placed in transverse 

 rows (not quite so regularly as shown in fig. 1 of PI. I.). As a rule, these spines are 

 narrow and pointed at the extremity which is attached to the wall of the body and 

 broadest at the other extremity. Here the free margin is deeply toothed, which gives 

 the spines a certain resemblance to the scales of the Lepidoptera. In other places the 

 incisions of the spines are so deep as to divide the scale into two or three narrow spines. A 

 small circular space at the peduncular pole is left free from spines, and at the other 

 extremity the terminal part is so completely covered with minute particles of mud and 

 sand, that it is impossible to distinguish the little spines there. This latter part of the 

 body is the only one which is visible when the little male is in its ordinary place, viz., 

 between the mantle or "sack" (as Darwin calls it) and the scutum of the hermaphrodite. 

 A small rounded part at the capitular extremity of the body is covered by a chitinous 

 membrane of greater thinness. The nuclei of the chitinogenous epithelium are placed 

 here much more closely and are more easily visible owing to the thinness of the 

 chitinous wall. A narrow slit-like opening (fig. l,o) divides this little circular space; it 

 corresponds with tlie orifice of the capitulum of the pedunculated Cirripedia. It is not 

 easy to distinguish the edges of this slit-like opening, owing, as Darwin suggested for the 

 same orifice of Scalpellum vulgare, to their extreme thinness. 



The chitinogenous membrane which is found beneath the chitinous outer wall shows 

 the ordinary structure of very flat cells with indistinct limits and with rather distant but 

 conspicuous nuclei. These nu;clei are very close to one another at the small circular 

 part at the capitular extremity (PI. III. figs. 2 and 3). The slit which indistinctly di^-ides 

 this part gives entrance to a cavity which contains the thoracic part of the little male. 

 This cavity is not lined by an epithelium ; it is only surrounded b)- a somewhat more 

 soUd layer of the same connective tissue, which fills up the whole interior of the 

 body of the male. This cavity is seen in transverse section in PL III. fig. 4. In all the 

 specimens of this species which I investigated the thoracic part was always retracted 

 high up into the interior of the body, so that even the verj'- long spines at the end 

 of the slender limbs never reached the slit-like orifice at the capitular pole. In the 

 males of some of the other sjaecies {Scalpellum intermedium, Scalpdlum tritonis) 

 the spines at the end of the thoracic limbs extend beyond this orifice. This was 

 often also the case in the males of Scalpellum vulgare as observed by Darwin, 

 which always showed the whole thorax forced outwards tkrough the orifice, a circum- 

 stance which perhaps was owing, according to Darwin, to the action of the spirits of 

 wine and consequent endosmose. 



Muscles of the hody-icall. —Under the cells of the hyiiodermis a well-developed layer 

 of muscular fibres is everywhere present ; these muscuhu- fibres are indistinctly trans- 

 versely striated ; in some of my preparations however the transverse striation is some- 



