384 BOTANICAL GAZETTE [MAY 
(44 per cent), but this obviously resulted from a loss of some pale 
individuals, which died off in early youth. The figure of 31 per 
cent was determined in June and should rather be compared with 
the percentages after self-fertilization determined at the flowering 
period (15-20 per cent). The Jorea seedling was evidently due to 
a mutation, even as after self-fertilization. 
O. grandiflora mut. lorea.—This mutant is characterized by its 
very narrow, almost linear foliage throughout its whole develop- 
ment. The leaves are dark green. The stature is almost the same 
as in the species, although at the end it is 2-3 dm. lower. Our 
climate, which is hardly favorable for the Alabama species, is still 
less so for this mutant. Not rarely the spikes miscarry, and bare 
anthers are of quite common occurrence. Especially in 1915 I 
found, during the whole summer, scarcely enough pollen for self- 
fertilization and some few crosses. The flowers are somewhat 
smaller and the petals less broad than in the parent species, and the 
fruits are thinner and more cylindrical. These differences are 
small, however, and probably a result of the insufficient nourish- 
ment by the narrow leaves. This latter character is always sharp 
and clear, and no intermediates have been observed. From two 
self-fertilized mutants of 1914 I cultivated a second generation, and 
from one of them I derived in 1916 the third one. They were uni- 
form lots and strikingly different from the original species. They 
embraced in 1915 in the first instance 60 specimens, all of which 
flowered, and in the second about roo seedlings, which were thrown 
away as soon as their uniformity was beyond doubt. The third 
generation in 1916 consisted of 61 plants, almost all of which 
flowered and resembled their parent. 
I crossed O. lorea with O. grandiflora in 1915, but could not find 
pollen for the reciprocal cross. In June 1916 I had among 59 indi- 
viduals 35 grandiflora, 15 ochracea, and 9 lorea, giving about 60, 25, 
and 15 per cent. The figure for ochracea is too low, since some 
seedlings were yellow and died in the seedpan, but it coincides 
sufficiently with the coefficient of mutation from the parent species 
as determined in the summer (15-20 per cent in table I). That for 
lorea is more reliable, since no losses could interfere here. It must 
be considered as due to the combination of all the mutated pollen 
