58 BOTANICAL GAZETTE [january 



Treub's figures of L. cernuum, was found in these specimens. 

 However, the base of the prothallium is more pointed than in 

 L. cernuum, and this pointed base may represent the primary 



tubercle. 



In proportion to the size of the prothallium, the protocorm is 

 much more massive than in L. cernuum. There is no single, 

 definitely organized growing point giving rise to all the protophylls, 

 but rather a series of points, each giving rise to a protophyll. 

 Stoma ta are abundant almost to the base of the protophyll; they 

 are of the simplest type and open into a loose parenchyma with 

 large air spaces. The transverse section is circular and shows a 

 single weak vascular strand extending a short distance into the 

 protocorm and ending blindly, without uniting with the strands 

 of neighboring protophylls. The protuberance shown in front of 

 the large protophyll in fig. 2 might be mistaken for the growing 

 point from which the leafy axis is to be developed, but that point 

 is formed much later, after several protophylls have appeared. 

 The prothallium and protocorm shown in fig. 2 are similar, but indi- 

 cate that there is considerable variation in both structures. Out- 

 lined against the protocorm is a second embryo. 



The much older protocorm (fig. 3) indicates that the protophylls 

 arise at irregular points, although there is a general progression, 

 so that the protocorm resembles a very short horizontal rhizome. 

 The 2 protophylls in the foreground are evidently the first ones 

 formed, and the 3 much smaller ones at the left are the latest. The 

 leafy axis of the permanent plant has not yet appeared. This 

 specimen and also those shown in figs. 1 and 2 were sectioned, but 

 the soil prevented satisfactory results. However, the sections 

 showed the position of sex organs, the distribution of the fungus, 

 and the relation of the protocorm to the prothallium. These 

 features are shown, in a very diagrammatic way, in fig. ib. 



L. later ale belongs definitely to the type represented by L. 

 cernuum and L. inundatum, since it has a short-lived green prothal- 

 lium and an ephemeral protocorm with protophylls preceding the 

 permanent leafy plant. L. salakense also belongs here, since the 

 prothallium is green, and in its earlier stages behaves like that of 

 L. cernuum; but it differs from the other 3 in having no endophytic 



