204 BOTANICAL GAZETTE [SEPTEMBER 
arrangement of the spiral of segments, as no exceptions were 
found in an examination of several other cases, although no exten- 
sive study of this feature was undertaken. 
The early apical cell forms a slightly compressed and slightly 
conical mass of cells. When the apical cell ceases to function, as 
in fig. 32, the embryo is more uniformly cylindrical, sometimes 
slightly club-shaped. The apical cell vanishes long before the stem 
tip, the cotyledons, or any of the body regions are recognizable, 
and nearly all of the early part of the embryo formed by apical cell 
growth goes to form the suspensor by the elongation of layer after 
layer of cells from the basal part of the embryo. 
ROSETTE AND ROSETTE EMBRYOS 
No investigator seems to have followed the development of 
the rosette further than through the early stages of elongation of 
the suspensor. That the open cells of the tier above the rosette 
disorganize has been stated by various workers. The writer has 
also been unable to find any traces of these nuclei of the upper open 
tier after the early stages of suspensor elongation, and doubtless 
they disintegrate. 
The rosette has usually been regarded as a group of cells between 
the main body of the egg and the suspensor, having no particular 
function. This view has proved to be erroneous, for the rosette 
is a group of young embryo initials which will produce embryos. 
These embryos are bounded by thick walls and are not so free to 
elongate as the primary embryos below them. 
After a little delay, during which the adjoining primary sus- 
pensor cells elongate, the rosette cells divide, as shown in one of the 
rosette cells of fig. 58, also in some of the rosette cells seen in polar 
view in fig. 59. A wall soon appears in one of the 2 daughter cells, 
inclined at an angle to the first (fig. 61, and rosette of fig. 46), 
forming the second segment of the apical cell. The apical cell 
continues to cut off segments on 2 or more sides, and the later 
embryo appears to have 3 cutting faces. Fig. 65 is a side view 
of a group of rosette embryos and shows well the apical cell and 
its segmentation, and (s) the upper portion of the collapsed primary 
suspensor. 
