1918] BUCHHOLZ—PINUS 205 
None of the rosette embryos has been found to reach stages much 
in advance of those shown in figs. 64-68. In some of these the 
embryonal tubes elongating from the basal portion of the embryo 
have formed a recognizable suspensor, which often appears freakish, 
as in fig. 68, modified no doubt by the unfavorable position and 
the unequal thickness of the walls of the rosette cells. 
It will be seen that the orientation of these rosette embryos 
is variable. In fig. 67 they have begun to elongate in various 
directions. The direction of the apical portion and the suspensor 
must be determined by the first few divisions, and figs. 59-64 
show that these are likewise quite variable. Before the rosette 
embryos have developed much beyond the early stages, such as 
fig. 59, the archegonium breaks down, and these embryos may be 
found pushed up against the top of the corrosion cavity by the 
suspensor. Even before the archegonium has completely broken 
down the rosette is frequently tilted by the twisting suspensor 
below, and it is quite probable that the orientation of the rosette 
embryos is related to the position of the rosette when the first divi- 
sions occur in these embryo initial cells, a thing that may well 
account for the lack of uniformity or regularity. 
It often happens that some of the rosette cells disorganize 
early and fail to produce embryos. Rosette cells may be found 
with no visible nuclei, or with nuclei in various stages of disinte- 
gration, while the neighboring rosette. cells are producing embryos. 
While these exceptions occur, it is evident that the normal product 
of an archegonium is 8 embryos. This makes polyembryony 
a much more extensive phenomenon than has hitherto been recog- 
nized. All of the species of Pinus investigated showed this peculi- 
arity, P. Banksiana, P. Laricio, P. echinata, and P. sylvestris. 
Rosette embryos develop less rapidly than the 4 primary embryos, 
abort in early stages, and it is entirely outside of the range of 
probability that they may ever contribute the embryo of the seed. 
ELONGATION OF THE ROSETTE.—Another abnormal phenomenon 
that was occasionally noted was that of elongated rosette cells 
resembling the primary suspensors. Fig. 53 shows a rosette in 
the first stages of elongation; fig. 54 shows another that is well 
advanced. Elongated rosette cells were found in nearly 5 per cent 
