292 BOTANICAL GAZETTE [OCTOBER 
of the spindle or in the development of the cell wall, since they 
disappear before the first division. However, they may represent 
an early assembling of the kinoplasmic substance preparatory to 
spindle formation. 
Fig. 6 shows the spirem partially segmented with the nucleolus _ 
near the center of the cavity and the segments projecting in from 
the nuclear membrane. Later the chromosomes become very 
short and group themselves about the nucleolus (fig. 7). At this 
time they suggest the tetrads described for many animals and 
closely resemble those of Arisaema triphyllum as figured by ATKIN- 
son (3). A typical bipolar spindle is formed and the bivalent 
chromosomes when arranged at the equatorial plate appear as 
very short X’s, V’s, and Y’s. 
Two cells separated by a distinct cell wall result. Son the 
heterotypic division and form the axial row (fig. 8). The micro- 
pylar cell of this row is smaller than the other, and disintegrates 
very quickly. The chalazal cell grows and is the mother of the 
embryo sac. An axial row of two cells is not common among the 
Archichlamydeae. TrEuB (43) describes an axial row of two cells 
for Viscum articulatum, and several cases have been reported 
among the monocotyledons. Miss Raitr (37) states that 4 
megaspores are formed in J. pallida, but makes no sketch showing 
them. In her fig. 1, J, she shows an ovule containing a large cell 
which she names the functional megaspore. Between it and the 
epidermis a small disintegrating cell appears. The sketch closely 
resembles the appearance of the ovules of J. Suliani with an axial . 
row of 2 cells and throws doubt upon her assertion that there is 
an axial row of 4 cells. 
The micropylar cell of the axial row is never large, and is so 
short-lived that it is easily overlooked, and the embryo sac seems 
to arise directly from the megaspore mother cell, as in Liliwm. 
As the micropylar cell disintegrates and the epidermal cells of the 
nucellus grow, there appears simply a small blue staining cavity 
between the embryo sac mother cell and the apical region of the 
epidermis, as cited by CouLTER and CHAMBERLAIN (15) for Clematis, 
and Helleborus (GUIGNARD 20), and Delphinium (MotTTIER 31). 
The chalazal cell grows and its nucleus divides, completing the 
