300 BOTANICAL GAZETTE [OCTOBER 
as it is wanting in the vicinity of the haustorium, which does 
not possess cell walls and would therefore appear in need of pro- 
tection. He thinks that the tapetal cells possess a ferment in 
their mucilaginous content and exercise a digestive function, for 
they persist while the neighboring tissue disintegrates, and they 
surround the parts which are in the process of rapid growth. 
Extending through the raphe is a strand of ceJls which is sur- 
rounded, except at the ends, by a layer of cells with cutinized walls. 
This strand terminates at the chalaza in the tissue of regular com- 
pact cells. It is into this that the antipodal region of the sac 
pushes (fig. 37). In looking at the figure it will be seen that the 
-outer layer of cutinized cells ends just as it passes this area, and 
on the inner side the layer ends almost in contact with the antipodal 
end of the tapetum. It thus forms a protective covering to the 
conducting tissue as it passes to the chalazal haustorium. No true 
vessels were ever observed in this strand of conductive cells. 
As noted earlier, the antipodal nuclei disappear before the 
haustorium develops, thereby giving rise to the unusual condition 
of a haustorium unaccompanied by nuclei. In none of the litera- 
ture studied was I able to find any record of a similar condition. 
In the cases where a haustorium has developed at the antipodal 
region before fertilization had occurred, it is usual for the antipodal 
nuclei to be present in the haustorium formed. An interesting 
example of this is given by SovEcEs (41) for the Solanaceae, where 
a pocket is formed at the basal part of the embryo sac and the 
antipodals take their place in the bottom of this and their digestive 
juices diffuse into the tissue beneath and dissolve out a cavity. The 
process of dissolution of the tissue varies among the different mem- 
bers of the family from a simple disjunction of the digestive layer 
of cells to a chalazal cavity whose capacity is comparable to that 
of the embryo sac itself, as in Lycopersicum esculentum. 
Stamen 
The flower possesses 5 stamens whose anthers are connivent and 
form a hood over the pistil. In a cross-section of a bud the sides 
of two adjacent anthers show as having their cells in contact, and 
this region appears as solid tissue. Each anther contains 4 micro- 
