308 BOTANICAL GAZETTE [OCTOBER 
formation of the secondary nucleus is constant for a given genus. 
In Lavatera and others it is formed before fertilization, while it is 
formed after in Hzbiscus. 
Following fertilization a micropylar haustorium is formed. 
The primary endosperm nucleus divides before the division of the 
fertilized egg. After a few divisions have taken place several of the 
resulting endosperm nuclei pass up through the micropylar part of 
the embryo sac (figs. 46, 47) out into the micropyle and form a 
haustorium which emerges from the micropyle and crosses over the 
space between the latter and the funiculus (figs. 49a, 6). As the 
haustorium encounters the funiculus it either extends along the 
funiculus some little distance before entering it or penetrates it 
immediately and branches freely within its tissue (fig. 53). 
In the early stages, as the endosperm nuclei pass through the 
upper part of the sac, they are surrounded by cytoplasm rendered 
dense by the presence of a large amount of food substance and 
consequently the nuclear membrane is entirely obscured (figs. 46, 
47). The densely staining filiform apparatus, all that remains of 
the two synergids, is still present, but it is more widely separated 
from the enlarged part of the embryo sac than in earlier stages 
(fig. 47). The narrow micropylar part of the sac where the syner- 
gids of the mature embryo sac were situated has widened and 
encroached upon the small disorganizing cells of the inner integu- 
ment. These cells have now disappeared except for a few remains 
and large regular cells limit the micropylar cavity into which the 
sac has pushed (cf. figs. 9, 14, 47).. At this stage the fertilized egg 
is still undivided and several endosperm nuclei lie below as well as 
above it. These nuclei are not shown in fig. 47, as they occurred 
in a different section from the one sketched. 
In the later stages in the development of the micropylar haus- 
torium but few nuclei are present in it. These are very large and 
contain a large nucleolus and stain bright red with safranin (fig. 53)- 
This haustorium was seen to persist up through the oldest stages 
studied, namely, those containing embryos with radicle and coty- 
ledons differentiated. 
The micropylar haustorium of J. Sultani differs from that of 
I. amphorata, as described by Lonco (28), in not entering the 
