1918] OTTLEY—IMPATIENS 309 
outer integument. It simply pierces the funiculus and branches 
extensively within it. 
As in I. amphorata (LONGO 28), I. Sultani possesses a chalazal 
haustorium as well as a micropylar one. This haustorium is much 
less extensive than is the micropylar one. One of the endosperm 
nuclei at the antipodal region becomes very large, and with its 
surrounding cytoplasm forms a long cell which pierces through the 
sac and one end of it enters the chalazal tissue, while the other end 
remains in contact with numerous normal endosperm cells (figs. 50, 
51, 53). 
It was difficult to secure a vertical section through this haus- 
torium, as a series of sections which cut through the embryo 
vertically would section the haustorium somewhat diagonally. 
Fig. 50 shows the outline of the cell, but no nucleus, while fig. 51 
shows the embryo sac portion with a large nucleus, but the chalazal 
part was cut so that all connection between chalazal tissue and 
haustorium was lost. This large haustorial cell contains a densely 
granular cytoplasm and is doubtless active in conveying nutriment 
from the chalaza to the endosperm. This haustorium does not 
remain active as long as the micropylar one. By the time the 
cotyledons of the embryo have become differentiated it is not so 
prominent as in earlier stages, while the micropylar haustorium 
still seems very active (fig. 53). 
The endosperm develops more rapidly at the micropylar and 
chalazal regions than at the sides of the sac. A layer of cytoplasm 
with a single row of free nuclei persists at the sides for some time, 
while at the poles of the sac walls come in early to separate the 
nuclei (figs. 49a, 50, 51). With the presence of two rows of nuclei 
at the sides, walls form about the outer layer of nuclei, while those 
of the inner layer remain free (fig. 48) until some time later, when 
they are separated by walls,and the endosperm is composed entirely 
of cells (fig. 53). The size and shape of the embryo sac have under- 
gone marked changes during endosperm formation. At the time 
of fertilization the outline of the sac is similar to that shown in 
fig. 54, with the early antipodal haustorium opposite the micropylar 
part of the sac. Later, during early endosperm formation, the sac 
elongates and widens slightly at the micropylar region. Below 
