1915] BURLINGAME—ARAUCARIA BRASILIENSIS 17 
lower part of the gametophyte enlarges many fold. This growth 
is due in part to cell multiplication, but more largely to the increase 
in size of the already existing cells. As they enlarge they form 
and store up starch and multiply their nuclei up to 4 or 5 in almost 
every cell, and in some of them to twice these numbers. Fig. 46 
shows nearly all of a single cell taken from about the middle of the 
endosperm. The light lines near the border mark the position of 
the delicate cell walls, which do not show in the photograph. The 
large oval bodies are starch grains, and the small round ones are 
proteids. Many of the cells are so crowded with food as to make 
photographs difficult. The proteid granules appear much later 
than the starch grains. They are not distinguishable optically 
much before the stage of the embryo shown in fig. 42. They never 
become so large or so numerous as the starch grains. Much the 
larger part of the growth of the gametophyte occurs during this 
period of food formation and storage subsequent to fertilization. 
At this period it is not more than 5-6 mm. in length, while at 
maturity it is about 4 cm. long and 15 mm. wide at the widest part. 
At fertilization it is broadest just below the archegonia; at 
maturity it is broadest at the basal end (compare fig. 48 with fig. 4 
in the earlier paper [3b]). 
After the embryo has differentiated its organs and has begun its 
final stage of development its cells become packed with food mate- 
rials (fig. 45). The smaller round grains shown in the figure are 
starch. The proteids occur in very large subspherical masses. 
Not infrequently the large globule includes a smaller one. The 
inclusions are also sometimes angular and probably crystalloids. 
The latter are smaller than the globular ones. Peculiar dumb- 
bell-shaped bodies are also found included in the large proteid 
masses. Often one end is included, while the other projects freely 
from the surface. The nuclei of these cells often become very 
large and sometimes flattened. Two conspicuous nuclei of this 
sort are shown along the lower side of fig. 4 
The growth of the gametophyte does sot destroy the nucellus, 
as usually happens among the gymnosperms. On the contrary, it 
continues to develop pari passu and forms an integral part of the 
mature seed coat. In fig. 48 it can easily be distinguished as a 
