1915] BURLINGAME—ARAUCARIA BRASILIENSIS 6 Fe 
3. There is some reason to suppose that some of the Cordaitales 
may have had ovules of the same general type as the lycopods just 
mentioned. If so, they were probably pollinated on the scale and 
might have given rise to modern conifers. 
4. It would be possible to derive modern conifers from a meso- 
zoic stock which had ovules and pollination apparatus comparable 
to that now possessed by the araucarians. 
Summary 
1. Pollination occurs on the scale at a distance from the 
nucellus. 
2. The pollen tube is very long and gives rise to many small 
lateral haustorial branches. It combines features of conifers and 
cycads to a certain extent. 
3. Reasons are adduced to show that this is probably an 
extremely primitive form of tube, having come down from very 
remote times little changed. 
4. The body cell divides in the extra-nucellar part of the tube 
a month or more before fertilization. The central cells of the 
archegonium divides very late or perhaps not at all, except in 
cases of delayed fertilization. 
5. The male cells are very large and unusually active, as well 
as long-lived. 
6. Blepharoplast-like bodies are found in the male cytoplasm. 
7- The male cells pass through the neck without injuring the cells. 
8. The male cell comes into violent contact with the egg and 
frequently displaces it. 
9. The free nuclear divisions of the proembryo are restricted 
to the male cytoplasm that surrounds the fusion nucleus, which 
persists and grows with the proembryo. 
to. The male cytoplasm around the older proembryo may be 
surrounded by a membrane. 
11. The number of free nuclei in the — varies from 32 to 
45 or perhaps more. 
12. When walls form the free nuclei are sities concentrically. 
13. The upper peripheral nuclei form the suspensor, the lower 
ones the cap, and the middle girdle elongates to unite cap and 
suspensor. 
