42 BOTANICAL GAZETTE [JANUARY 
male and female plants have well marked characters. The an- 
theridial heads are decidedly globose and show variations in color 
from yellow brown to red brown, occasionally almost black. The 
archegonial heads are less globose, have a somewhat flattened 
aspect on top, and show no unusual coloring except the conspicuous 
bud at the growing point in the center of the head. This bud varies 
in color from yellow brown to red brown. An analysis of the bud 
reveals archegonia almost mature on short side branches near the 
apex of the main axis, the coloring matter being in the perichaetial 
leaves surrounding the organs. 
A careful study of the whole bog and others for several miles 
about disclosed the fact that not a single sterile head of S. sub- 
secundum could be found. Sex organs were everywhere in vast 
numbers. In order to determine whether or not such a condition 
might be unusual, and to provide abundant material for study, 
developments through winter, spring, summer, and autumn of 
1913 were closely followed; and again in the autumn of 1913 the 
sex organs appeared in the same vast numbers. 
As far as S. subsecundum in the vicinity of Mineral Springs is 
concerned, we conclude therefore (1) that sex organs are not of rare 
occurrence; (2) that both antheridial and archegonial heads on 
approaching maturity are distinctly characterized by coloration; 
(3) that antheridia and archegonia begin to develop in August and 
September respectively, the antheridia always appearing first. 
A brief statement of time relations in the developmental process 
of the archegonium as observed in the autumn of 1913 may be 
of interest. Young stages were first noted on September 16 and 
continued to appear for approximately four weeks. By October 25 
the youngest archegonia were beginning the formation of canal 
cells, while the oldest were almost mature. At this time the 
coloring of the perichaetial leaves began to be noticeable. By 
November 15 the canal row had broken down in some of the oldest 
archegonia. Those archegonia which have not reached maturity 
on the advent of cold weather develop slowly through the winter. 
In the spring, therefore, at the time of the disappearance of the 
snow, it is possible to find stages having 7 or 8 canal cells, with the 
ventral cell not yet divided. 
