46 BOTANICAL GAZETTE [JANUARY 
Hence figs. 6-9, belonging to this latter class, are known to be 
transverse. 
THE SECONDARY ARCHEGONIUM 
The secondary archegonium shows a greater degree of uniform- 
ity. The initial divides into an inner and an outer cell (figs. 6, 
10). This outer cell by subsequent transverse divisions (figs. 8, 9, 
14) gives rise to a filament of cells, 5 or 6 in number, in each of 
which the usual secondary divisions occur (fig. 160). As yet no 
evidence has been found that the secondary archegonium may 
develop by an apical cell with two cutting faces. 
THE DEVELOPMENT OF THE ARCHEGONIUM PROPER 
After there has been formed, as described above, a filament 
of cells by transverse walls, or a series of cells by an apical cell with 
two cutting faces, and secondary divisions have occurred in each 
segment except the terminal one, the development of the arche- 
gonium proper begins in the manner usual among the Bryophytes. 
In the terminal cell, which becomes somewhat enlarged, three 
oblique walls appear, cutting off three peripheral segments and 
originating a large cell within, which has the form of an inverted 
truncated pyramid (figs. 13, 14). This large cell we shall designate 
the primary axial cell. On division it gives rise to an outer axial 
cell, the cover cell, and an inner axial cell, the central cell (figs. 
17-19). The wall cells of the archegonium arise from the three 
peripheral segments. 
Up to this point the development of the archegonium proper 
coincides exactly with the description given by numerous investi- 
gators for the archegonium of the Bryophytes, whether Hepaticae 
or Musci. It is in the events immediately following that there is 
a divergence of views and theories. For the sake of continuity we 
shall postpone the presentation of these theories until later and 
continue the description of the developmental processes. 
Here then the important question arises, What is the part 
played by the cover cell and what by the central cell ip the further 
development of the archegonium? The answer must be found in 
a study of sections both longitudinal and transverse. 
