52 BOTANICAL GAZETTE [JANUARY 
Summary 
1. During the autumns of 1912 and 1913 sex organs have been 
found in vast numbers on Sphagnum subsecundum in the vicinity 
of Mineral Springs, Indiana. 
2. On approaching maturity the archegonial heads may be 
recognized by the colored bud in the center of the head at the apex. 
Analysis of the bud shows terminal archegonia on short side 
branches. 
3. The archegonia begin to develop in September. 
4. The apical cell of a side branch is a primordium; each of 
the two segments last formed becomes the initial of a secondary 
archegonium, while that part of the apical cell above and not 
included by these segments is the initial of the primary archegonium. 
5. There is great irregularity in the early stages of the develop- | 
ment of the primary archegonium: there may be a filament of 
cells by the successive transverse divisions of the apical cell; or 
growth by an apical cell with two cutting faces; or a mixture of 
planes. 
6. As yet the secondary archegonium has been found to develop 
only by the successive transverse divisions of the apical cell. 
7. The archegonium proper is initiated in the manner usual 
among the Bryophytes. In the terminal cells three oblique walls 
cut off three peripheral segments and originate the primary axial 
cell within, which on division gives rise to cover cell and central 
cell. 
8. The cover cell is relatively inactive and cuts off no basal 
segments. 
- g. The central cell on division forms the primary neck canal cell 
(the mother cell of the neck canal row) and the primary ventral 
cell. . 
10. The growth of the neck canal row is intercalary, the cells 
dividing in almost any order. 
11. The primary ventral cell divides late into ventral canal 
cell and egg. 
12. The growth of the wall cells of the archegonium is inter- 
calary. 
13. The mature archegonium has 8 or g canal cells. 
