126 BOTANICAL GAZETTE [FEBRUARY 
In general behavior this embryo sac differs in no way from the 
rest of the Scrophulariaceae, in which it is a general rule that the 
synergids are well marked off from the other nuclei of the embryo 
sac, and the antipodals are inconspicuous, disappearing about the 
time of maturity. 
Case of an ovule with two embryo sacs 
In an ovary in which most of the ovules were in the 4-nucleate 
or 8-nucleate stages, one ovule was seen showing two sacs lying 
side by side. Fig. 18 represents a section through this ovule not 
passing through the embryo sacs. In this section the ovule has 
every appearance of having been derived from two fused ovules. 
Figs. 16 and 17 support this view, as the cells separating the two 
embryo sacs clearly belong to the integument. Unfortunately the 
sections through this ovule are oblique. Figs. 13-17 are drawn 
from this ovule as it was represented in five consecutive sections. 
The left-hand embryo sac has four nuclei arranged in two groups 
of two each. The right-hand embryo sac contains eight nuclei 
apparently not definitely arranged in groups. Six of the nuclei lie 
fairly close together at one end of the sac and two at the other end. 
Possibly this lack of arrangement is to be correlated with the abnor- 
mal conditions under which the sac has developed. This ovule was 
the only one showing any abnormal tendencies. 
Cases in which two ovules have grown together to form one with 
two embryo sacs have not been reported as frequently as those in 
which the two embryo sacs arise from a single archesporium. They 
have been recorded, however, in Pirus Malus, Loranthus europaeus, 
and Viscum album (5). 
Fertilization 
Ovules in which fertilization is taking place are fairly abundant. 
In the section represented in fig. 12 the pollen tube is seen having 
penetrated the embryo sac and apparently pushed its way through 
one of the synergids, and has discharged its contents into the sac. 
One male nucleus is seen fusing with the egg, while the other is 
fusing with the nucleus produced by the fusion of the polar nuclei. 
The male nuclei may be easily distinguished from the nuclei of the 
