156 BOTANICAL GAZETTE [FEBRUARY 
appear shortly after the first ones. In the mature plant the whole 
inflorescence and the upper part of the stem is glandular, but no 
glands appear at all on the staminate flowers. 
The paths of the axial bundles are the same as those in the 
staminate flowers, with two exceptions: first, the strand in the 
center of the whorl of five strands at the base of the ovarian cavity 
is not abortive, but passes into the ovule and curves with the 
integument to a point level with the antipodal cells (fig. 20, a); 
second, even when there are stamens no branch of the upper corolla 
strand branches into them. After fertilization, the style and 
corolla wither and drop off. When this happens, the whorl of five 
strands which lies adjacent to the integument (fig. 20, b) disorgan- 
izes and disappears, along with tissues which are also adjacent to 
the integument, thus freeing the ovary from the integument. 
The style is cleft (fig. 20, st?) and the stigma is covered with 
papillae like those on the staminate flowers. 
When the ovarian cavity begins to develop, the growth of the 
stamens is retarded. Usually they disappear, but sometimes 
remain as seen in fig. 20, st, but none bearing pollen sacs were seen. 
There was nothing unusual found in the formation of pollen 
grains. The outer wall layer becomes thick and spiny, while the 
spore becomes winged. In grooves between the wings, the extine 
is merely in contact. At the three germinative spots the walls are 
pushed outward (fig. 23). This occurs before the pollen grains 
leave the anther. 
Summary 
The development of the staminate and pistillate flowers is the 
same up to the development of the ovarian cavity. 
Both staminate and pistillate flowers have ovarian cavities, 
but ovules develop only in the pistillate flowers. 
Stamens are sometimes found in pistillate flowers, but they are 
always sterile. 
No gland-hairs are found on staminate flowers. 
Styles of staminate flowers are cleft. 
Nothing unusual occurs in the development of the egg, embryo, 
or pollen grains. 
UNIVERSITY OF WASHINGTON 
SEATTLE, WASH. 
