1915] PETRY—OPHIOGLOSSACEAE 349 
immediately below. LANG (g) has given in detail the vascular 
connections of the branches in five specimens of this species; he 
concluded that ‘‘the chief vascular supply of the branch is derived 
from a development of xylem adaxially to the subtending leaf trace.” 
This adaxial xylem is considered to be an extension of the margins 
of the leaf trace, and therefore centrifugal in character. 
Botrychium lanceolatum var. angustisegmentum.—A single 
branching specimen of this species was secured. The terminal 
bud has been destroyed and the branch has arisen at a considerable 
distance below. As shown by 
fig. 2, the vascular system of 
the branch connects with the 
subtending leaf trace. After 
the trace has been separated 
from the stem stele for a con- 
siderable distance, tracheids 
appear on the adaxial side of 
the protoxylem of the trace; as 
shown by fig. 3, these are 
definitely centripetal in origin 
and occur in contact with the 
Protoxylem. The mass of 
ag es Saas aces Fic. 3-—Detail of leaf trace of fig. 2, B: 
, e, endodermis; 157. 
soon assumes the characters of 
the main stele. This differs from the general condition in B. 
Lunaria, as described by LANG, in that the branch connection 
arises from centripetal xylem of the leaf trace rather than from 
adaxial extension of the centrifugal xylem. 
Botrychium ramosum.—Two rhizomes of this species, each bear- 
ing a single branch, were secured. In each case the terminal bud 
has been destroyed and the branch has developed at a considerable 
distance below. The vascular connections of the branches differ 
greatly from that described above. 
Fig. 4 shows the vascular supply of the branch in the first of 
these specimens. As the leaf trace swings away from the stem 
Stele, wings of primary and secondary xylem develop in the angle 
WX 
Sy 
Be $s 
