1915] BERRY—MYRTACEAE 485 
America, and these include the only monotypic genera in the sub- 
family (Orthostemon Berg, Psidiopsis Berg, and Paivaea Berg), as 
well as the large and greatly differentiated genera like Myrcia DC. 
with about 450 existing species. The two other large genera, 
Myrtus Linn. with about 178 existing species, and Eugenia Linn. 
with about 1300 existing species, are the only two genera found on 
all the continents (except Europe), and in these two genera America 
furnishes 135 species of Myrtus and about 850 species of Eugenia, 
or over 75 per cent in Myrtus and over 65 per cent in Eugenia. 
The second subfamily, the Leptospermoideae, comprises the 
Leptospermae with 28 genera and about 700 existing species, and 
the Chamaelaucieae with 12 genera and about 165 existing species. 
Both of these tribes are even more strikingly Australian than the 
Myrtoideae are American. The Chamaelaucieae are entirely 
Australian and are mainly confined to Western Australia, and in 
accordance with their peculiar habitat, specialized characters, 
and restricted range, are probably of relatively recent origin. The 
Leptospermae have a single monotypic genus in Chile, and the dis- 
tribution of the other members of this tribe suggests the probability 
that the South American genus should be placed in some other 
alliance, since with the exception of Metrosideros Banks, which is 
represented in Africa, and the genus Baeckea Linn., which reaches 
the Asiatic mainland, all of the genera are confined to Australia 
or to the surrounding islands southeast of Asia. 
ANDREWS,” in a recent paper, has presented some interesting 
Statistics of distribution and an ingenious theory of the history 
of the family. He considers that the original stock was arborescent 
or shrubby, with entire, simple, opposite, penni-veined leaves with 
dots and intramarginal acrodrome veins; with the calyx lobes and 
petals imbricate, probably in fives; flowers regular, solitary or in 
cymes; stamens indefinite, numerous, free, with versatile, 2-celled 
anthers; ovary inferior, with two or more cells; style simple; 
fruit inferior, crowned with persistent limb of calyx, indehiscent, 
succulent or fleshy (rarely dry); albumen none; cotyledons thick 
and fleshy, with a short radicle. 
2 AnDREws, E. C., The development of the natural order Myrtaceae. Proc. 
Linn. Soc. N.S. Wales 38:5297568. 1913. 
