8 BOTANICAL GAZETTE [JULY 
make it certain that the growth of both canal row and peripheral 
cells of the neck is intercalary and not apical. 
SUMMARY 
In regard to the formation of the archegonium proper in the 
Musci 3 theories have been advanced: (1) the HOFMEISTER con- 
ception of the tangential division of one of the 4 original pedicel 
rows, a theory soon made untenable by the work of later investi- 
gators; (2) a recent revival of the HorMEISTER scheme modified 
by the tangential division of one segment only, as proposed by SER- 
VETTAZ; (3) the commonly accepted account, confirmed again and 
again for all the great groups of the Bryophyta, namely, the appear- 
ance in the terminal cell of 3 oblique walls forming 3 peripheral 
segments and an axial cell within. 
It cannot be maintained, therefore, that the origin of the 
archegonium proper is in doubt. The evidence is too overwhelm- 
ing to admit of any uncertainty on this point. However, the 
development of the axial row is another matter. Here is a subject 
involving widely conflicting accounts, some being diametrically 
opposed. Summarized, these accounts are: (1) the abandoned 
conception of HorMEISTER, having only a historic interest; (2) 
Kitun’s claim for Andreaea that all the cells of the axial row are 
cut from the base of the apical cell; (3) CAMPBELL states for Funaria 
_ that the axial row is composed of a primary canal cell and seg- 
‘ments cut from the base of the apical cell, none of which divide — 
after they are formed, so far as could be determined; (4) GorBEL 
holds that in Mnium undulatum the topmost neck canal cell (the 
‘one just below the cover cell) acts as an apical cell in the produc- — 
- tion of the canal row; (5) Janczewskt finds that the cells of the 
axial row are of diverse origins, the upper arising through trans- — 
_ verse divisions of the 2-6 initials cut from the base of the apical 
cell, while the lower are formed by the t jons of the 
inal initial; © Hotrerty has shown ‘that the growth in : 
- the canal row of Afni um Ch o 
(7) Gavet concludes that, the canal cells among the Musci have 
S all the s same ae : that there are no segments cut from the oe : 
