10 BOTANICAL GAZETTE [JULY 
for Funaria, but extend to the basal walls. The primary axial cell, 
therefore, has something of the shape of an inverted, truncated 
pyramid. There now follows the division of the primary axial 
cell into an outer axial cell, the cover cell, and an inner axial cell, 
the central cell (figs. 12, 13). Quite soon the central cell divides. 
the resulting lower cell being the ventral cell, while the upper 
is the primary neck canal cell (figs. 14, 15). The actual division 
of the central cell was found twice. The axial row of the young 
archegonium now consists of the ventral cell; its sister cell, the 
primary neck canal cell; and a large cover cell (figs. 15-17). : 
It is interesting to note that in fig. 15 the original division wall 
between cover cell and central cell appears tilted, due partly to the 
inequality in the growth of the peripheral segments, and partly to 
the change in the direction of the axis through the formation of 
new peripheral segments by the apical cell. While this tilting is — 
not always found, it is of frequent occurrence, as shown to a greater 
or less extent in figs. 16, 18, 20, and furnishes valuable evidence in 
separating that portion of the axial row derived from the central 
cell from the part contributed by the cover. 
The cover cell now cuts off peripheral segments ns 15-17). 
No absolute Proof can be given as to their exact number, but it 
series as figs.15-18 that there __ 
are e3 peripheral segments. Then there is added to the canal row 
an initial cut from the base of the cover cell. The evidence for 
this statement rests on fig. 18, on several others quite like it, and __ 
: is corroborated by the figures in the series about it (figs. 16-20). 
similar series could be constructed from the material — 
oe Sears A Jong and careful search failed to reveal the actual | 
division figure, but in fig. 18 the size and position of the nuclei and _ 
2 z the delicate wall between leave no doubt that the uppermost canal 
|) ow consiéts of the ventral cell; its sister cell, the primary r 
a >I] has t cut from t base of the cover cell and that the: process a 
has just: been completed. As illustrated by fig. 19 the axial Tow 
canal cell; an initial cut from the base of the cover cell; and a large ae 
: . cover cell or apical cell. 
Up to this point the | process of development i is ae and deh 
ease Meee tee ability shockir 
