t9t7] DUPLER—TAXUS CANADENSIS 121 
The body cell soon migrates into the tube and is followed by 
the stalk cell nucleus, the two usually being in contact and appar- 
ently having passed into the tube together. During the growth 
of the tube through the nucellus, up to the time when the pollen 
tube reaches the female gametophyte, the 3 nuclei are in an axial 
row, while later the stalk nucleus migrates around the body cell and 
es a position besides the tube nucleus. Of these two nuclei in 
the tube, the tube nucleus is generally to be recognized by its being 
slightly larger than the stalk nucleus. The body cell begins enlarg- 
ing at once after entering the tube. As it passes down the tube it 
is slightly ellipsoid, but on reaching the end of the tube, as the 
latter is enlarging about the endosperm, it becomes rounded as it 
increases in size. The cytoplasm remains dense and the nucleus 
becomes large. During the early history of the body cell its cyto- 
plasm is uniformly Soneented through the cell, but : as it reaches 
Teo wake pet vn laonn Kaenrrow Lass 
Ps 
and here eae delicate radial strands (fig. 23). "The nucleus, 
which earlier occupied a central position, takes a more peripheral 
one (usually on the side opposite to that on which the stalk and 
tube nuclei lie), while the cytoplasm becomes considerably vacuo- 
lated along the margin of the cell, he eRe Sinner peering 
_ with the increased vacuolization. . 
The division of the body cell into the two male cells takes place : 
shortly before fertilization. Several mitot n 
were found. — _ The nucleus of the body cell being at one side of the 
_ cell, there is an unequal division of the cytoplasmic material. ae 
_A broad spindle is formed and the cell plate laid down on it is 2 
the ¢ 
lenticular i in outline, resulting i in the. formati 
