1917] DUPLER—TAXUS CANADENSIS 129 
two gametophytes, only the archegonia of the lower gametophyte, 
however, being fertilized 
In my material I find two gametophytes quite common, usually 
in an axial row, although sometimes lying side by side (fig. 57). 
One is usually larger than the other, the upper generally being the 
smaller of the two, and both may produce archegonia, as pointed 
out by Coker. Usually when 2 gametophytes develop in an 
axial row the pollen tubes push in between them, and then the 
archegonia are directed toward the tubes; but this is not always 
the case, as sometimes there are 2 apparently equally vigorous 
gametophytes, one above the other, and both with good archegonia 
in their micropylar ends. The pollen tube has spread out above the 
upper gametophyte, while the lower one is not in contact with the 
tube at any point in the case shown in fig. 58. 
Several instances of more than 2 gametophytes in an wre were 
found. One case (fig. 59) shows 3 gametophytes with tissue, the 
pollen tube lying between the two uppermost. No archegonia 
were present in the upper one, but both of the lower ones have good 
oe. pie of which has beer: fertilized and contains a pro- 
embryo. Other cases show hytes, in in one of which (fig. 60) 
2 of the gametophytes have formed tissue, while the other 2 are in 
free nuclear condition, archegonia being present only in the lowest, 
which also contains a proembryo, the pollen tube in this: Pag a 
having pushed its way between the 4 gametophytes. — In 
- 
case (fig. 61) the ovule contains 3 g with tissue. ee 
archegonia, while the fourth one consists of only a few paeaart 3 
and, although lying between two ‘vigorous gametophytes with 
tissue, retains its shape, evidently owing to the firmness of the 
megaspore membrane. One ovule was ; found containing 5 ere ae 
