THE ANATOMY OF SIX EPIPHYTIC SPECIES OF 



LYCOPODIUM 



CONTRIBUTIONS FROM THE HULL BOTANICAL LABORATORY 1 89 



J. Ben Hill 

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(with twenty-eight figures) 



In this investigation, six epiphytic species of Lycopodium were 

 studied. Two New Zealand species, L. Billardieri Spring, and 

 L. varium R. Br., with two South African species, L. verticillatum 

 G.f. and L. Holstii Hieron., were furnished by Dr. Charles J. 

 Chamberlain; and two forms from Samoa, Z. Phlegmaria Linn, 

 and L. carinatum Desv., were collected by Dr. W. J. G. Land. 

 The two species from New Zealand were identified by Dr. Jesse M. 

 Greenman of the Field Museum, Chicago, and the remainder were 

 identified by Dr. G. Hieronymus of Berlin. There was some doubt 

 concerning the identity of the two forms from Samoa, since they 

 do not correspond exactly to the descriptions of their respective 

 species. Dr. Hieronymus suggests that each represents a new 

 variety or subspecies. The species studied are described as grow- 

 ing epiphytic upon trees, with the short, vegetative branches 

 more or less erect, and the long, slender, dichotomously branched 

 strobili pendulous (fig. i). Most of these species are rather rare, 

 but L. Phlegmaria and L. Billardieri are mentioned in the recent 

 literature dealing with the morphology of the group. Since these 

 forms are not very familiar to residents of the northern hemisphere, 

 a short general description may serve to make this account more 



intelligible. 



L. Holstii is a very coarse plant, with short thick stems covered 

 with large stiff leaves. The stem is 4-6 mm. in diameter, with a 

 small stele 0.35 mm. in diameter. The leaves are about 2.5 cm. 

 long and sharply pointed. Type material, furnished by Dr. G. 

 Hieronymus, shows sporophylls with little or no modification 

 from the vegetative leaves. They are not grouped in a definite 

 strobilus, but are simply aggregated near the apex, much as in 

 L. lucidulum. 



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[Botanical Gazette, vol. 58 



