150 BOTANICAL GAZETTE [august 



in Polygonal urn, from 2 in Smilacina racemosa, S. amplexicaulis, 

 and Streptopus roseus, and from 4 in Smilacina stellata, S. sessifolia, 

 Maianthemiim canadense, and Medeola virginica. 



2. In all forms in which more than one reduction nucleus enters 

 into the structure of the embryo sac, these nuclei are at first more 

 or less completely separated by cell membranes, the degree of 

 separation varying from split cell plates in Smilacina stellata to 

 evanescent cell plates in Medeola virginica. 



3. The difference in the degree of the separation of these cells 

 cannot affect their morphological status; they are all megaspores. 



4. In the light of this evidence it seems reasonable to conclude 

 that all reduction nuclei arising from the nucleus of the megaspore 

 mother cell, whether temporarily separated or not separated at all, 

 should be regarded as megaspore nuclei. 



5. Two megaspore mother cells were occasionally observed in 

 six of the eight species investigated. 



University of Texas 



Austin^ Texas 



LITERATURE CITED 



1. 



Bernard, C. H., Recherches sur les spheres attractives chez Lilium candi- 



diinij etc. Jour. Botanique 14:118-124, 177-188, 206-212. 1900. 

 2. Brittox, Nathaniel Lord, Manual of the flora of the northern states 



and Canada. New York. 1901. 

 3- Brown, William H., The nature of the embryo sac of Peperomia. Bot. 



Gaz. 46:445-460. 1908. 



4- ' — , The embryo sac of Habenaria. 



5. Brown, William H., and Sharp, Lester W., The embryo sac of Epi- 

 pactis. Box. Gaz. 52:439-452. 191 1. 



6. Campbell, D. H., The embryo sac of Pandanus. Bull. Torr. Bot. Club 

 36:205-220. 1909. 



7- , The embryo sac of Paftdanus. Ann. Botany 25:773-789. 1911. 



Bot. Gaz. 48:241-250. 1909 



W 



wild oat, Avena fatua. Proc. 

 9. Chamberlain, C. J., Winter 



Gaz. 25:124-128. 1898. 

 10. Coulter, J. M., Relation of 

 Bot. Gaz. 45-361-366. 1908. 



900 



Bot. 



J. M., and Chamberlain, C. J., Morphology 



New York. 1903. 



