356 BOTAXICAL GAZETTE [October 



Under ordinary conditions sporangiola of the type just described 

 are formed almost to the exclusion of other forms of sporangia; 

 but, especially when the culture is made in a saturated atmosphere, 

 typical sporangia appear, associated with them, having the normal 

 structure of the ordinary sporangia of the Mucoraceae, and similar 

 to those which occur in Choanephora. These sporangia, however, 

 are subject to very great variation, and almost every imaginable 

 intermediate condition between a form such as is represented in 

 fig. 1 6, and the three-spored sporangiolum may be met with. These 

 single sporangia when well developed, as in the instance just re- 

 ferred to, possess a typical columella, and contain very numerous 

 spores, which are more uniform and usually smaller than such as 

 occur in smaller sporangia. Their characters are in general similar 

 to those of the spores formed in the sporangiola; they possess the 

 same appendages, which are perhaps slightly longer and are similarly 

 though less distinctly marked. Their color is often more reddish 

 brown, but is variable. They may rarely be borne almost erect, but 

 are usually nodding or strongly circinate, the termination of the 

 sporangiophore in many cases being coiled in a more or less irregular 

 spiral. Such large forms with small spores are far less frequent 

 than smaller sporangia which show the greatest variability in size 

 (figs. 13-15 and 17), some of which may be hardly larger than the 

 typical sporangiola and like them may possess no columella (fig. 14) • 

 On the other hand, abnormalities or more simple conditions of the 

 sporangiolum type of fructification occur which, like that illustrated 

 in fig. 12, serve to break down any clean-cut distinction between 

 the two types. The spores of the smaller sporangia are likely to be 

 larger than those of better developed examples (figs. i^ 1 S)> anc * 

 the walls are more coarsely roughened. The wall of the sporangio- 

 phore, and even sometimes of hyphae which bear sporangiola, 

 may be roughened by a scaly incrustation, sometimes confined 

 to the summit, sometimes extending to the base. 



Chlamydospores are produced usually in enormous numbers, 

 especially under unfavorable conditions, and vary considerably in 

 form, from cylindrical to nearly spherical. 



Although no "conidia" appear to be produced by this species, 

 it is evident that it must be regarded as very closely related to 



