1912] SHULL— LYCHNIS AND PAP AVER 129 



from the usual type, the margin being in this case red-violet 6 instead 

 of nearly white. Whether this red-violet margin was a purely 

 somatic modification of the dark-red body-color, or whether it 

 was germinal, it was clearly of a different nature from the white 

 margins involved in the other families. 



In the two families (10287, 10289) whose margined parents 

 were evidently homozygous, a small number of plants were recorded 

 without margins. These exceptional plants occurred among those 

 set into the garden, while larger numbers of plants from the same 

 families, which were grown to maturity in pots in the propagating- 

 house, were all margined. In family 10287 there were 6 plants 

 with unmargined petals among 40 grown to maturity in the garden, 

 and none among 133 which flowered in pots, and in family 10289 

 one was noted as unmargined among 47 plants in the garden and 

 none among 83 which developed in pots. However these seven 

 unmargined specimens are to be accounted for, it is clear that 

 each of these families is the offspring of a homozygous margined 

 parent. 



In the 25 ma tings between plants, neither of which possessed 

 margined petals, there appeared only 15 plants with margins 

 among a total of 1402 offspring, and in a number of those recorded 

 as margined the margin was merely a trace of lighter color of more 

 or less doubtful character. Only in one family (10291) were the 

 margins unmistakable, and in this family the margined plants 

 occurred only among those which were retained in the greenhouse. 

 Of 21 which matured in the garden none had margins, while among 

 99 which flowered in pots in the greenhouse there were 10 with 

 margins, several having only a trace, while others had well marked 

 white margins 2 mm. wide — in one plant 3 mm. wide. No attempt 

 need be made at present to account for these few margined plants, 

 for their number is too small to vitiate the conclusion that the 

 unmargined condition is recessive, and that typically all the 

 offspring of two unmargined parents are unmargined. 



The most interesting matings in which margins were involved 



6 The color-nomenclature adopted in this paper is based on the spectrum colors, 

 as arranged in the Milton Bradley system. Exact shades and tints have been recorded, 

 but for the sake of simplicity these have not been reproduced here. 



