TENDRILS OF SMILAX 1 



Agnes Arber 



(with plate xxii) 



more 



almost 



vegetative morphology of flowering plants. It is unnecessary to 

 deal here in detail with the voluminous literature which has grown 

 up about the subject, since this task has been excellently performed 



himself 



structures 



to be well founded. It will suffice to group together the various 



enumerated, and to attempt 



test them, from 



typical 



The views 



follows, under live heads : 



imul gated may 



metamorphosed 



outgrowths of the apical region of the leaf-sheath. 



Both these possibilities seem 



immediately above the sheath 

 from it both externally and as 



their 



2. That the Smilax tendrils represent a bipartite ligule, each 

 tendril being a demi-ligule. 



This alternative is ruled out because, as Gluck (5) and Domin 

 (4) have shown, there is occasionally a definite ligule in addition 

 to the tendrils. It is also discounted for the same reasons that 



make the first view untenable. 



3- Th 

 segments 



metam 



lateral free nerves which 



the 



1 This paper represents part of the work carried out during the tenure of a Keddey 

 Fletcher-Warr Studentship of the University of London, and with the aid of a 

 grant from the Dixon Fund of the University of London. The writer wishes to 

 acknowledge her indebtedness for material to Miss G. Lister, Professor Seward, and 

 the Curator of the Cambridge Botanic Garden. 



Botanical Gazette, vol. 69] 



[438 



