1913] BURLINGAME—ARAUCARIA BRASILIENSIS IIL 
is strengthened by the occurrence of more than two sperms in 
Microcycas (3) and Ceratozamia (4). The deposition of the pollen 
at a distance from the nucellus has also been recorded for Saxe- 
gothaea cons picua (15), and serves to strengthen the likeness of the 
araucarians and podocarps. 
By long odds the most interesting and suggestive feature of the 
male gametophyte of the araucarians yet known is the remarkably 
large ‘‘sperms”’ and the possibility of their motility. The one 
figured (plate fig. 29) is about 150 long. It is by no means the 
largest one observed, but seemed to be the one most definitely 
organized. Such dimensions are only known elsewhere among the 
actively swimming sperms of the Cycadales and Gingko. It is of 
interest to note that the kauri (6) appears to have two large nuclei 
also, though it is not stated that they constitute part of organized 
cells. Inasmuch as Agathis has usually been held to be more 
primitive in most respects than Araucaria, it may be possible that 
they will yet be found there. The aster-like fibers found in the 
dividing body cell are of course very suggestive of a blepharoplast, 
and one might perhaps legitimately expect such structures to be 
found in connection with such large and possibly motile male cells. 
Again they might be only such structures as have been recorded by 
LAND (12) in the egg of Ephedra. 
Summary 
1. The staminate cones are extraordinarily large and have 
numerous sporophylls with an indefinite number of pollen sacs 
pendent from the abaxial side of the swollen apex. 
2. An almost incomprehensibly great number of pollen grains 
is produced. 
3. The method of differentiating the sporogenous tissue is 
variable and indefinite. The size of the sporangium and the 
number of microspores is subject to wide fluctuations. 
4. The structures concerned in dehiscence are very fernlike. 
5. The chromosome number in the male gametophyte is 8. 
6. Prothallial tissue is formed in a manner almost identical with 
that in the Podocarpineae, but the number of cells so formed is 
greater. 
