Reena arr tahoe ay ot ja, de eed Wee apt eee 22 ysl Se ee 
NESE teraes Soe eer ame coer 5 oy peer ecg eee a ee Te 
r a Borie 
1913] CHAMBERLAIN—MACROZAMIA MOOREI 147 
Among the oriental cycads (Cycas, Macrozamia, Bowenia, 
Encephalartos, and Stangeria) the situation is not so uniform. In 
Cycas the ovulate plant does not bear a compact cone, the sporo- 
phylls having the position of a crown of vegetative leaves with the 
growing point at the center, the growing point persisting from the 
seedling throughout the life of the plant. In the staminate plant 
there is hardly ever more than one cone, and this is terminal. 
There are cone domes in the pith. ° 
In Bowenia, both ovulate and staminate cones are borne singly 
on slender branches of the main stem. 
In Stangeria, both ovulate and staminate cones are terminal and 
are usually borne singly, although occasionally there may be two 
or three, especially in case of staminate plants. There are well 
marked cone domes in the pith. 
In Encephalartos, both male and female plants often bear more 
than one cone, and in some species a single cone is the exception, 
while three, four, and five are common. PEARSON noted that in 
Encephalartos Frederici Gulielmi the cones are arranged in a circle 
about the bud. In this species, at Queenstown, South Africa, 
where PEARSON made his observation, I saw seven ovulate cones in 
a circle about a well developed bud. In E. caffer, E. Altensteinii, 
E. horridus, and E. villosus I found three to five cones in a circle 
about a well developed bud. Such cones are lateral with respect 
to the growing point, which does not become transformed into a 
cone, but continues the growth of the plant. A dissection of adult 
specimens of E. Altensteinii and E. villosus showed no cone domes 
in the pith, and cone domes are necessarily present when cones are 
terminal. It may be doubtful whether cones are terminal in 
Encephalartos, even when produced singly. 
In various species of Macrozamia ovulate plants frequently 
bear more than one cone, and in staminate plants more than one 
cone is the rule. Both ovulate and staminate cones are obviously 
lateral, even when borne singly. There are no cone domes in the 
pith. 
Of course, in all the genera, when there is branching in the 
popular sense of the term, each branch may bear a cone. 
Since Macrozamia Moorei approaches so closely to the Ben- 
