WIDDRINGTONIA IN SOUTH TROPICAL AFRICA. 191 
the diameter of the canal varies in different leaves, and in different 
parts of the same leaf; in the broader parts of the leaf two smaller 
canals are often present, one on each side of the median. Below the 
epidermis, especially in the middle line of the ey is a band of 
mechanical tissue, and transfusion tracheids ore or less 
plentiful to the right and left of the vascular paridllde A similar 
arrangement of parts occurs in the South African W. cupressoides, 
and also in W. juniperoides, from the Cedarberg Mountains. In my 
original song ede of W. Whytet (Trans. Linn. Soc. ser. 2, iv. 
, t. ix. figs. 7 & 8) Pibitradtal ond leaf-anatomy of W, juni- 
peroides and W. Whytet from this point of view; examination of 
a larger series of specimens shows that in W. juniperotdes two 
lateral canals are sometimes present. 
There is some variation in the size of the cones, but this varia- 
tion is shown in specimens from Mt. Milangi, which are admittedly 
conspecific. Thus the specimens from Umtali bear smaller cones 
closely resembling those on a specimen from Milangi at Kew, col- 
lected by Mr. D. B. Ritcher; this Milangi specimen also bears 
larger more robust cones, similar to those of the original Milangi 
specimens from Mr. Whyte. The cones of the Rhodesian speci- 
mens, so far as I have seen, never r reach the size or robustness of 
the larger Milangi specimens. The cone-scales have a similar 
texture in all the “specimens, showing a rough somewhat tubereled 
dorsal surface, different = from the extremely tubercled scales of 
W. juniperoides and the smooth-backed scales of W. cupressotdes. 
The cones of the two South African ae es are ai Paps 
S 
mens, 
re ver, insufficient to justify a 
specific separation, and the evidence points to the existence of one 
species, namely, tien a Whytei, common to this area of 
South-east Tropical A 
r. Selater is stoned that local “oon ascribes the intro- 
duction of the Umtali specimens to the Queen of Sheba. They are 
wn locally as cedars, and were sieimmadts the descendants of 
—_ ig shea Pacey seeds brought back by the Queen from her visit 
King 
a. 
rary 
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id 
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,2 
— 
In the geabaeacen of the leaf-sections, a large series of which has 
been examined, I have to acknowledge the help of Mr. W. Williams, 
Demonstrator in Botany at the Birkbeck College. 
EXPLANATION OF PhatE 479B. 
Transverse sections of leaves of Widdringtonia Whytei, from Umtali, 
Rhodesia » x90. 
1. Showin wing Sasa and lateral resin-canals. 2. Showing single median 
resin-canal. m, t —— mechanical tissue; 7, resin-canal ; tr, transfusion 
tissue ; v. D. eda | bun 
