6 BOTANICAL GAZETTE [JANUARY 
Farther within (P, fig. 19) is another bell-shaped area which 
takes a deeper stain than the fundamental tissue on either side. 
This is the fundament of the inner portion of the pileus. 
The remaining primordial tissue lying between the fundament 
of the pileus and that of the stipe is in the form of a hollow cone, 
the wall of which is broad at its base. It is the tissue which 
DEBary (3, p. 204) called the cone (Kegel). In the stained sec- 
tion, a photograph of which is represented in figs. 18, 19, this cone 
of primordial tissue does not show a homogeneous structure. At I 
there is a deeply stained area, also campanulate in form when taken 
as a whole, lying within the lighter colored portion of the cone. 
This is what I regard as the fundament of a true indusium, which 
in Ithyphallus impudicus does not become further differentiated, 
so far as present evidence indicates, though the rudiment may 
persist in a recognizable form (in magnified section) up to the 
maturity of the plant. The hyphae here are somewhat more 
densely interwoven, and therefore this zone stains more deeply 
in contrast with the other primordial tissue on either side of it. 
This rudiment of an indusium described by FiscHER (6, p. 26) 
remains in the condition of primordial tissue, which, with some 
adjacent primordial tissue, in the mature plant forms a thin 
membrane lying between the pileus and the stipe. My inter- 
pretation of the fundaments represented by these more deeply 
staining cones in the young fruit body of IJthyphallus impu- 
dicus is slightly different from the interpretation given by FISCHER. 
The zones in fig. 19 indicated by P (pileus fundament), J (indusium 
fundament), and B (primordial tissue lying between) are considered 
by FiscHER as one zone, the fundament of an indusium which he 
marks in his figures as J. An examination of his figures shows 
that it is made up of three layers or zones, two outer ones more 
deeply stained than the inner one, corresponding to the three zones 
of my figure (P, B, and J). Fiscuer states that the outer layer 
of his zone J forms the inner layer of the pileus, and that the 
indusium in Dictyophora phalloidea Desvaux is differentiated along 
the inner layer of the zone J. Zone J of my figure joins the stem 
at the same place as the inner layer of Fiscter’s zone J, and also 
in the mature fruit body, as will be shown later, the zone J of my 
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