26 BOTANICAL GAZETTE [JANUARY 
going paragraphs, that our specimen cannot belong in any way to 
the genus as at present anatomically characterized. For in the 
very important particulars of wood structure, organization of the — 
phloem, as well as the constitution of the pith and cortex, it differs 
absolutely from all existing members of the Cupressineae and a 
Taxodineae. It is equally clear, from a consideration of all the 
features of histological structure just indicated, that the cone 
axis under discussion belongs with those Araucarineae of the more : 
primitive type which became extinct with the close of the Mesozoic — 
period. There remains, however, one point to be elucidated. It 
has been shown in the memoir on the conifers of Kreischerville, 
frequently cited above, that the vegetative twigs known as Sequoia 
Reichenbachit had structure of the Brachyoxylon type, that is, a 
Paracedroxylon type described by Sinnott. Our cone, 
beyond any reasonable question belongs in the same genus, has 
not the Brachyoxylon type of wood, but on the contrary shows all 
the features of ligneous organization found in Paracedroxylon. — 
at the present time, which is applicable to an even wider range 
anatomical facts, that the reproductive axes of this order of g 
sperms are apt to perpetuate the structure of ancestral. types. 
‘For example, the structure of the wood in the cones of existin| 
pines is now known to retain the type of wood structure foun 
the vegetative branches of the pines of the Lower Cretaceous. 
Applying this principle to the cone axis at present under consider- 
ation, it is rendered probable that the Paracedroxylon type was 
characteristic of araucarian vegetative twigs representing the 
ancestral forms from which the so-called cretaceous sequoias were 
derived. We have thus another and independent argument 
the stock, but that the Brachyoxylon, Araucariopitys, and Pores 
cedroxylon araucarian types are all older than Araucarioxylon and 
connect it in one way or another with the type of wood found in the 
