94 BOTANICAL GAZETTE [FEBRUARY 
of the two that prevail generally. As the pericycle is delicate and 
irregular, the protoxylem of the root often abuts on the endodermis. 
In a few specimens spiral exarch protoxylem is present in the 
cotyledonary node. This is continuous with the protoxylem at the 
poles of the primary root and in the first leaf trace. In other 
stems the mesarch position characteristic of the older stem appears 
at the cotyledonary node. The same unsettled condition persists 
in the solenostele, although an examination of the apical region 
shows that the first elements lignified are toward the center of 
the stem. In the strands of the dictyostele definite protoxylem 
points appear in an endarch position, with occasionally one or two 
centripetal elements beyond them (fig. 22). These bundles do not — 
show the distinction between large bundles with mesarch or no 
protoxylem, and small strands with endarch protoxylem observed 
in Angiopteris. 
The cotyledonary trace shows the same kind of variations in the 
position of the protoxylem as the cotyledonary node. It may be 
exarch where it joins the stele (fig. 26) and gradually shift to 
mesarch (figs. 26-30) in the petiole; no protoxylem may be dis- 
tinguishable in the trace near the stele, but a development of it 
in a mesarch position occurs while the trace is passing through the 
cortex; or the trace may consist of only two or three scattered 
elements. The next three or four leaves are mesarch, and may 
become endarch in the upper part of their course. The older 
leaf traces are endarch as in Angiopleris and Danaea. 
APICAL MERISTEMS 
The apical regions of Marattia have been the source of much 
difference of opinion. Hormetster (13) described a single deep 
triangular apical cell for the Marattiaceae, as for all other vascular 
cryptogams. According to HoLtE (14) a four-sided apical cell 1s 
found in the stem of the young sporophyte and is retained pet 
manently. Bower (2) describes for mature plants a 4 or 5-celled 
meristem meeting at the intersection of two more or less perpe?- 
dicular lines. CAMPBELL (4) found that in some cases at least 
the apex of the stem is occupied by a single initial. FARME® 
and Hitt (9) describe an apical cell of irregular prismatic form, 
