tgtt] HAGU E—DIOSPY ROS 35 
able that perfect flowers are borne some seasons, as has been re- 
ported from Kansas." 
The staminate flowers are smaller than the pistillate and in 
clusters (fig. 1), 16 fertile stamens surrounding the sterile pistil. 
The pistillate flowers are solitary, and usually contain 8 sterile 
stamens, but very often the number is greater. 
The early stages of the development of the two flowers are the 
same. The floral cycles are generally preceded by a pair of bracts, 
though often there is only one (fig. 2). The calyx next appears 
(fig. 3) and becomes a massive enveloping cup before the other 
cycles can be seen, which appear in centripetal sequence (fig. 4). 
The corolla can be distinguished before the stamens, but they 
develop together in the typical sympetalous fashion. Occasionally 
the calyx or corolla has more than four parts; this is illustrated 
in fig. 6, in which the calyx has five divisions. 
The stamens of the staminate flower fork (fig. 5), thus doubling 
the pistillate number (fig. 6); in the pistillate flower it is a common 
occurrence to find the number increased by the branching of one 
or more of the stamens. The fertile pistil contains eight ovules. 
The style is single, but the stigma is four-parted. Not many 
sterile pistils were examined; those that were had no ovules and 
a short imperfect style. 
Megasporangium and megaspores 
The ovule is anatropous and has two integuments (fig. 7), this 
last character being unusual among the Sympetalae. The mother 
cell can be distinguished by its size and conspicuous nucleus about 
the time the inner integument is first visible (fig. 8). Judging by 
the repeated appearance of this stage in the material, it is espe- 
cially persistent. Only one mother cell occurs in a sporangium, and 
is always next to the outer layer of nucellar cells, no parietal cell 
being cut off. One complete figure of the first division of the 
mother cell was found in the spindle stage (fig. 9). Compared 
with the preceding conspicuous nucleus of the mother cell, the 
spindle is small and has very small and numerous chromosomes. 
* The Industrialist, Kansas State Agricultural College, March 1904. 
