36 BOTANICAL GAZETTE [JULY 
A portion of another figure showing the formation of the wall 
between the daughter cells was seen. 
Four megaspores are formed in a linear row (figs. ro, 11), but 
it is not always complete, three cells not being uncommon. One 
exception to the usual arrangement was discovered, in which the 
outer daughter cell is divided by a vertical wall (fig. 12). As 
usual, the chalazal megaspore becomes the embryo sac (fig. 13). 
Early stages of the embryo sac 
The two and four-celled stages of the embryo sac were not 
found. At the eight-celled stage the sac is small, much longer 
than wide, and somewhat pointed at the micropylar end. When 
the sac is seen enveloped by the single nucellar layer, it appears 
decidedly cone-shaped, and is supported upon a stalklike portion 
of the nucellus as upon a pedestal (fig. 15). The growth of the 
sac does not obliterate this nucellar tissue until a comparatively 
late stage of the ovule. In the sac of fig. 14, which is the eight- 
celled stage in which the polars are differentiated, only seven 
nuclei are shown, and it seems quite probable that that is the full 
number for that particular sac, because there is much evidence 
that the usual number of nuclei is not always present. This con- 
clusion is reached because of the conspicuous absence of antipo- 
dals. Three antipodals were found in one sac, but only after 
a long search. Extremely early disintegration would also account 
for the absence of these cells, but no evidence was found for this 
explanation. 
The egg apparatus in the eight-celled stage shows nothing 
unusual. The three cells are in the ordinary position, and there 
is the customary differentiation of the cells in size. 
In striking contrast to these two groups, the antipodals and 
egg apparatus, are the polars, which are large and conspicuous 
(figs. 21a, 216), and are found either approaching or fusing in 
material gathered during the flowering time. 
During the development of the sac the integuments become 
massive, and the innermost layer of cells of the inner integument 
becomes large and full of protoplasm, forming a tapetal layer com- 
