rgtt] BROW N—LACHNEA SCUTELLATA 281 
decrease in the size of the nuclei during the development of the asco- 
carp and the persistence of the same number of chromosomes 
throughout the ascogonium and ascogenous hyphae, moreover, 
indicate very strongly that a fusion of nuclei during this stage is 
not to be expected. 
When the ascogonium has reached its mature size, it gives off 
a number of large ascogenous hyphae which are multinucleate 
from the first (plate fig. 3). The nuclei do not appear to be 
arranged in pairs or in any 
other definite manner, but 
to be scattered irregularly 
in the hyphae (fig. 14). 
They are undergoing divi- 
sion rather rapidly, as has 
been previously described. 
About this time the cyto- 
plasm and nuclei of the 
other cells of the archicarp 
begin to degenerate. These 
cells apparently do not fuse 
together as in Ascophanus 
carneus (CUTTING 7). The 
ascogenous hyphae grow 
up among the vegetative 
hyphae which are situated Fics. 14-16.—Fig. 14, outgrowth of ascoge 
over the ascogonium and _ nous hyphae from ascogonium; fig. 15, storage 
have been mentioned as ‘lls giving off paraphyses; fig. 16, tips of 
aE ‘ ascogenous hyphae in hymenium; all X 525. 
giving rise to paraphyses. 
As the ascogenous hyphae increase in length, they branch freely 
and become divided up into a number of large multinucleate cells. 
Some nuclei are left in the ascogonium and these finally degenerate. 
When the ascogenous hyphae are growing out from the ascogonium, 
the vegetative cells over the ascogonium (plate fig. 3) are slender, 
densely protoplasmic, and extend upward toward the covering of 
the ascocarp. They thus have the appearance of young paraphyses, 
but do not take part in the formation of the hymenium until they 
have developed further. As they grow up they branch freely and 
